馬尾杉屬

马尾杉属
	覆葉馬尾杉
科学分类
界：	植物界 Plantae
演化支：	维管植物 Tracheophyta
纲：	石松綱 Lycopodiopsida
目：	石松目 Lycopodiales
科：	石松科 Lycopodiaceae
属：	马尾杉属 Phlegmariurus; (Herter) Holub, 1964
种
	见正文

馬尾杉屬（学名：Phlegmariurus）为石松目下的一个屬。馬尾杉屬下植物的孢子囊穗外觀呈繩索狀，且向下懸垂，看來好比馬尾，因此得名。

属名Phlegmariurus源于希腊语phlegmaria，意为“粘液质的”。

特質

石松目植物的歷史可追溯到古生代，結構與最早的維管束植物相似，具有分叉的莖，鱗狀葉小且只有1條葉脈，葉子基部的孢子囊有同形孢子^[1]。距今3億4千萬年前的石炭紀初期曾有高度超過40公尺的石松類植物出現^[1]。

分類問題

過去植物分類幾乎把所有石松類植物都歸入石松屬下^[2]。目前趨勢則是分成幾個不同的屬，因此很多馬尾杉屬植物也被稱為石松，通常叫做XX石松的植物就是XX馬尾杉，例如覆葉石松就是覆葉馬尾杉，展葉馬尾杉也有展葉石松的別名^[2]。

下属物种

本属包括以下物种：^[3]

Phlegmariurus acerosus (Sw.) B.Øllg.
Phlegmariurus acifolius (Rolleri) B.Øllg.
Phlegmariurus acutus (Rolleri) B.Øllg.
Phlegmariurus affinis (Trevis.) B.Øllg.
Phlegmariurus afromontanus (Pic.Serm.) A.R.Field & Bostock
Phlegmariurus afromontanus Pic.Serm.
Phlegmariurus ambrensis (Rakotondr.) A.R.Field & Bauret
Phlegmariurus amentaceus (B.Øllg.) B.Øllg.
Phlegmariurus andinus (Rosenst.) B.Øllg.
Phlegmariurus apolinarimarii (Nessel)
Phlegmariurus aqualupianus (Spring) B.Øllg.
Phlegmariurus arcuatus (B.Øllg.) B.Øllg.
Phlegmariurus ascendens (Nessel) B.Øllg.
Phlegmariurus attenuatus (Spring) B.Øllg.
Phlegmariurus australis (Willd.) A.R.Field
Phlegmariurus austroecuadoricus (B.Øllg.) B.Øllg.
华南马尾杉 Phlegmariurus austrosinicus (Ching) Li Bing Zhang
Phlegmariurus axillaris (Roxb.)
Phlegmariurus badinianus (B.Øllg. & P.G.Windisch) B.Øllg.
Phlegmariurus balansae (Herter) A.R.Field & Bostock
Phlegmariurus bampsianus (Pic.Serm.) A.R.Field & Bostock
Phlegmariurus banayanicus (Herter) A.R.Field & Bostock
Phlegmariurus beitelii (B.Øllg.) B.Øllg.
Phlegmariurus biformis (Hook.) B.Øllg.
Phlegmariurus billardierei (Spring) Trevis.
Phlegmariurus binervius (Herter) B.Øllg.
Phlegmariurus bolanicus (Rosenst.) A.R.Field & Bostock
Phlegmariurus brachiatus (Maxon) B.Øllg.
Phlegmariurus brachystachys (Baker) A.R.Field & Bostock
Phlegmariurus bradeorum (Christ) B.Øllg.
Phlegmariurus brevifolius (Hook. & Grev.) B.Øllg.
Phlegmariurus brongniartii (Spring) B.Øllg.
Phlegmariurus buesii (Herter) B.Øllg.
Phlegmariurus callitrichifolius (Mett.) B.Øllg.
Phlegmariurus campianus (B.Øllg.) B.Øllg.
网络马尾杉 Phlegmariurus cancellatus (Spring) Ching
Phlegmariurus capellae (Herter) B.Øllg.
Phlegmariurus capillaris (Sodiro) B.Øllg.
龙骨马尾杉 Phlegmariurus carinatus (Poir.) Ching
Phlegmariurus catacachiensis (Nessel) B.Øllg.
Phlegmariurus cavifolius (C.Chr.) A.R.Field & Bostock
Phlegmariurus chamaeleon (Herter) B.Øllg.
长叶马尾杉 Phlegmariurus changii T.Y.Hsieh
Phlegmariurus chiricanus (Maxon) B.Øllg.
Phlegmariurus christii (Silveira) B.Øllg.
Phlegmariurus ciliatospiculatus B.Øllg.
Phlegmariurus ciliolatus (B.Øllg.) B.Øllg.
Phlegmariurus cleefianus (B.Øllg.) B.Øllg.
Phlegmariurus cocuyensis B.Øllg.^[4]
Phlegmariurus colanensis (B.Øllg.) B.Øllg.
Phlegmariurus columnaris (B.Øllg.) B.Øllg.
Phlegmariurus comans (Herter ex Nessel) B.Øllg.
Phlegmariurus compactus (Hook.) B.Øllg.
Phlegmariurus copelandianus (R.C.Y.Chou & Bartlett) A.R.Field
Phlegmariurus coralium (Spring) A.R.Field & Bostock
Phlegmariurus costaricensis (Herter)
Phlegmariurus crassus (Humb. & Bonpl. ex Willd.) B.Øllg.
Phlegmariurus creber (Alderw.) A.R.Field & Bostock
Phlegmariurus crucis-australis (Herter) B.Øllg.
Phlegmariurus cruentus
(Spring) B.Øllg.
- Phlegmariurus cruentus var. exilis^[4]
- Phlegmariurus cruentus var. illimitatus^[4]
柳杉叶马尾杉 Phlegmariurus cryptomerianus ^[5]
Phlegmariurus cryptomerinus (Maxim.) Satou
Phlegmariurus cuernavacensis (Underw. & F.E.Lloyd) B.Øllg.
Phlegmariurus cumingii (Nessel) B.Øllg.
Phlegmariurus cuneifolius (Hieron.) B.Øllg.
杉形马尾杉 Phlegmariurus cunninghamioides (Hayata) Ching
Phlegmariurus curiosus (Herter) A.R.Field & Bostock
Phlegmariurus curvifolius (Kunze) B.Øllg.
Phlegmariurus dacrydioides (Baker) A.R.Field & Bostock
Phlegmariurus dalhousieanus (Spring) A.R.Field & Bostock
Phlegmariurus darwinianus (Herter ex Nessel) B.Øllg.
Phlegmariurus delbrueckii (Herter) A.R.Field & Bostock
Phlegmariurus deminuens (Herter) B.Øllg.
Phlegmariurus dentatus (Herter) Arana
Phlegmariurus dianae (Herter) B.Øllg.
Phlegmariurus dichaeoides (Maxon) B.Øllg.
Phlegmariurus dichotomus (Jacq.) W.H.Wagner
Phlegmariurus dielsii (Herter) A.R.Field & Bostock
Phlegmariurus divergens (Alderw.) A.R.Field & Testo
Phlegmariurus durus (Pic.Serm.) A.R.Field & Bostock
Phlegmariurus echinatus (Spring) B.Øllg.
Phlegmariurus ellenbeckii (Nessel) A.R.Field & Bostock
Phlegmariurus elmeri (Herter) A.R.Field & Bostock
Phlegmariurus engleri (Hieron. & Herter) B.Øllg.
Phlegmariurus eremorum (Rolleri) B.Øllg.
Phlegmariurus ericifolius (C.Presl) B.Øllg.
Phlegmariurus erythrocaulos (Fée) B.Øllg.
Phlegmariurus espinosanus (B.Øllg.) B.Øllg.
Phlegmariurus eversus (Poir.) B.Øllg.
金丝条马尾杉 Phlegmariurus fargesii (Herter) Ching
Phlegmariurus filicaulos (Copel.) A.R.Field & Testo
Phlegmariurus filiformis (Sw.) W.H.Wagner
Phlegmariurus firmus (Mett.) B.Øllg.
Phlegmariurus flagellaceus (Kuhn) A.R.Field & Bostock
Phlegmariurus flexibilis (Fée) B.Øllg.
Phlegmariurus foliaceus (Maxon) B.Øllg.
Phlegmariurus foliosus (Copel.) A.R.Field & Bostock
Phlegmariurus fontinaloides (Spring) B.Øllg.
福氏马尾杉 Phlegmariurus fordii (Baker) Ching
Phlegmariurus friburgensis (Herter ex Nessel & Hoehne) B.Øllg.
Phlegmariurus funiculus (Herter)
Phlegmariurus funiformis (Cham. ex Spring) B.Øllg.
Phlegmariurus gagnepainianus (Herter) A.R.Field & Bostock
Phlegmariurus galapagensis (O.J.Hamann) B.Øllg.
Phlegmariurus giganteus (Herter) A.R.Field & Bostock
Phlegmariurus gnidioides (L.fil.) A.R.Field & Bostock
Phlegmariurus goebelii (Nessel) A.R.Field & Bostock
Phlegmariurus gracilis A.Rojas & R.Calderón
广东马尾杉 Phlegmariurus guandongensis Ching
Phlegmariurus gunturensis (Alderw.) A.R.Field & Bostock
Phlegmariurus haeckelii (Herter) A.R.Field & Testo
Phlegmariurus haitensis (Herter)
喜马拉雅马尾杉 Phlegmariurus hamiltonii (Spreng.) Á.Löve & D.Löve
Phlegmariurus harmsii (Nessel) A.R.Field & Bostock
Phlegmariurus hartwegianus (Spring) B.Øllg.
Phlegmariurus hastatus (B.Øllg.) B.Øllg.
Phlegmariurus hellwigii (Warb.) A.R.Field & Bostock
Phlegmariurus hemleri (Nessel) B.Øllg.
椭圆马尾杉 Phlegmariurus henryi (Baker) Ching，椭圆叶马尾杉
Phlegmariurus heterocarpos (Fée) B.Øllg.
Phlegmariurus heteroclitus (Desv. ex Poir.) B.Øllg.
Phlegmariurus hexastichus (B.Øllg. & P.G.Windisch) B.Øllg.
Phlegmariurus hildebrandtii (Herter) A.R.Field & Bauret
Phlegmariurus hippurideus (Christ) B.Øllg.
Phlegmariurus hippuris (Desv. ex Poir.) A.R.Field & Testo
Phlegmariurus hoffmannii (Maxon) B.Øllg.
Phlegmariurus hohenackeri (Herter) B.Øllg.
Phlegmariurus holstii (Hieron.) A.R.Field & Bostock
Phlegmariurus homocarpus (Herter) B.Øllg.
Phlegmariurus horizontalis (Nessel) A.R.Field & Bostock
Phlegmariurus huberi (B.Øllg.) B.Øllg.
Phlegmariurus hugoi B.Øllg.
Phlegmariurus humbertii (Nessel) A.R.Field & Bostock
Phlegmariurus humbertii-henrici (Herter) A.R.Field & Bostock
Phlegmariurus hypogaeus (B.Øllg.) B.Øllg.
Phlegmariurus hystrix (Herter) B.Øllg.
Phlegmariurus idroboi B.Øllg.^[4]
Phlegmariurus iminii Kiew
Phlegmariurus insularis (Carmich.)
Phlegmariurus intermedius (Trevis.) B.Øllg.
Phlegmariurus itambensis (B.Øllg. & P.G.Windisch) B.Øllg.
Phlegmariurus jaegeri (Herter) A.R.Field & Bostock
Phlegmariurus josesantae B.Øllg.^[4]
Phlegmariurus kajewskii (Copel.) A.R.Field & Testo
Phlegmariurus killipii (Herter) B.Øllg.
Phlegmariurus kuesteri (Nessel) B.Øllg.
Phlegmariurus kwangtungensis C.Y.Yang
Phlegmariurus lancifolius (Maxon) B.Øllg.
Phlegmariurus lauterbachii (E.Pritz. ex K.Schum. & Lauterb.) A.R.Field & Bostock
Phlegmariurus lecomteanus (Nessel) A.R.Field & Bostock
Phlegmariurus ledermannii (Herter) A.R.Field & Bostock
Phlegmariurus lehnertii Testo
Phlegmariurus leptodon (Maxon ex Herter)
Phlegmariurus lignosus (Herter) B.Øllg.
Phlegmariurus lindenii (Spring) B.Øllg.
Phlegmariurus linifolius (L.) B.Øllg.
Phlegmariurus llanganatensis (B.Øllg.) B.Øllg.
Phlegmariurus lockyeri (D.L.Jones & B.Gray) A.R.Field & Bostock
Phlegmariurus loefgrenianus (Silveira) B.Øllg.
Phlegmariurus longus (Copel.) A.R.Field & Bostock
Phlegmariurus loxensis (B.Øllg.) B.Øllg.
Phlegmariurus luteynii B.Øllg.
Phlegmariurus macbridei (Herter) B.Øllg.
Phlegmariurus macgregorii (Baker) A.R.Field & Bostock
Phlegmariurus macrostachys (Hook. ex Spring) N.C.Nair & S.R.Ghosh
Phlegmariurus magnusianus (Herter) A.R.Field & Testo
Phlegmariurus mandiocanus (Raddi) B.Øllg.
Phlegmariurus mannii (Hillebr.) W.H.Wagner
Phlegmariurus marsupiiformis (D.L.Jones & B.Gray) A.R.Field & Bostock
Phlegmariurus martii (Wawra) B.Øllg.
Phlegmariurus megastachyus (Baker) A.R.Field & Bostock
Phlegmariurus melanesicus (Brownlie) A.R.Field & Testo
Phlegmariurus merrillii (Herter) A.R.Field & Bostock
Phlegmariurus mesoamericanus (B.Øllg.) B.Øllg.
墨西哥马尾杉 Phlegmariurus mexicanus (Herter) B.Øllg.
Phlegmariurus milbraedii (Herter) A.R.Field & Bostock, 2013
Phlegmariurus mildbraedii (Herter) A.R.Field & Bostock
闽浙马尾杉 Phlegmariurus mingcheensis Ching
Phlegmariurus minutifolius (Alderw.) A.R.Field & Bostock
Phlegmariurus mirabilis (Willd.) A.R.Field & Testo
Phlegmariurus mollicomus (Spring) B.Øllg.
Phlegmariurus monticola Kiew
Phlegmariurus mooreanus (Sander ex Baker) B.Øllg.
Phlegmariurus multifarius (Alderw.) A.R.Field & Bostock
Phlegmariurus myrsinites (Lam.) B.Øllg.
Phlegmariurus myrtifolius (G.Forst.) A.R.Field & Bostock
Phlegmariurus myrtuosus (Spring) B.Øllg.
Phlegmariurus neocaledonicus (Nessel) A.R.Field & Bostock
Phlegmariurus nesselii (Brause ex Nessel) B.Øllg.
Phlegmariurus niligaricus (Spring) A.R.Field & Bostock
Phlegmariurus nudus (Christ ex Nessel & Hoehne) B.Øllg.
Phlegmariurus nummularifolius (Blume) Ching
Phlegmariurus nutans (Brack.) W.H.Wagner
聂拉木马尾杉 Phlegmariurus nyalamensis (Ching & S.K.Wu) H.S.Kung & L.B.Zhang，也拼作Phlegmariurus nylamensis
Phlegmariurus obovalifolius (Bonap.) A.R.Field & Testo
Phlegmariurus obtusifolius (P.Beauv.) A.R.Field & Bostock
Phlegmariurus ocananus (Herter) B.Øllg.
Phlegmariurus oceanianus (Herter) A.R.Field & Bostock
Phlegmariurus oellgaardii (A.Rojas) B.Øllg.
Phlegmariurus oltmannsii (Herter ex Nessel) A.R.Field & Bostock
Phlegmariurus ophioglossoides (Lam.) A.R.Field & Bostock
Phlegmariurus orizabae (Underw. & F.E.Lloyd) B.Øllg.
卵叶马尾杉 Phlegmariurus ovatifolius (Ching) W.M.Chu ex H.S.Kung & Li Bing Zhang
Phlegmariurus pachyphyllus (Kuhn ex Herter) A.R.Field & Testo
Phlegmariurus pachyskelos B.Øllg.
Phlegmariurus parksii (Copel.) A.R.Field & Bostock
Phlegmariurus patentissimus (Alderw.) A.R.Field & Bostock
Phlegmariurus pecten (Baker) A.R.Field & Bostock
Phlegmariurus perrierianus (Tardieu) A.R.Field & Bostock
Phlegmariurus pflanzii (Nessel) B.Øllg.
马尾杉 Phlegmariurus phlegmaria (L.) Holub
Phlegmariurus phlegmarioides (Gaudich.) A.R.Field & Bostock
Phlegmariurus phylicifolius (Desv. ex Poir.) B.Øllg.
Phlegmariurus phyllanthus (Hook. & Arn.) R.D.Dixit
Phlegmariurus picardae (Christ ex Krug) B.Øllg.
Phlegmariurus pichianus (Tardieu) A.R.Field & Bostock
Phlegmariurus pinifolius (Trevis.) Kiew
Phlegmariurus pithyoides (Schltdl. & Cham.) B.Øllg.
Phlegmariurus pittieri (Christ) B.Øllg.
Phlegmariurus podocarpensis B.Øllg.
Phlegmariurus polycarpos (Kunze) B.Øllg.
Phlegmariurus polycladus (Sodiro)
Phlegmariurus polydactylus (B.Øllg.) B.Øllg.
Phlegmariurus polylepidetorum (B.Øllg.) B.Øllg.
Phlegmariurus portoricensis (Underw. & F.E.Lloyd)
Phlegmariurus pringlei (Underw. & F.E.Lloyd) B.Øllg.
Phlegmariurus proliferus (Blume) A.R.Field & Bostock
Phlegmariurus pruinosus (Hieron. & Herter) B.Øllg.
Phlegmariurus pseudovarius (Brownlie) Rouhan & A.R.Field
美丽马尾杉 Phlegmariurus pulcherrimus (Hook. & Grev.) Á.Löve & D.Löve
Phlegmariurus pungentifolius (Silveira) B.Øllg.
Phlegmariurus quadrifariatus (Bory) B.Øllg.
Phlegmariurus recurvifolius (Rolleri) B.Øllg.
Phlegmariurus reflexus (Lam.) B.Øllg.
Phlegmariurus regnellii (Maxon) B.Øllg.
Phlegmariurus ribourtii (Herter) A.R.Field & Bostock
Phlegmariurus riobambensis (Nessel) B.Øllg.
Phlegmariurus robustus (Klotzsch) B.Øllg.
Phlegmariurus rosenstockianus (Herter) B.Øllg.
Phlegmariurus rostrifolius (Silveira) B.Øllg.
Phlegmariurus ruber (Cham. & Schltdl.) B.Øllg.
Phlegmariurus rubricus (Herter) A.R.Field & Bostock
Phlegmariurus rufescens (Hook.) B.Øllg.
Phlegmariurus rupicola (Alderw.) A.R.Field & Bostock
Phlegmariurus sagasteguianus (B.Øllg.) B.Øllg.
柔软马尾杉 Phlegmariurus salvinioides (Herter) Ching
Phlegmariurus samoanus (Herter ex Nessel) A.R.Field & Bostock
Phlegmariurus sarmentosus (Spring) B.Øllg.
Phlegmariurus saururus (Lam.) B.Øllg.
Phlegmariurus scabridus (B.Øllg.) B.Øllg.
Phlegmariurus schlechteri (E.Pritz.) A.R.Field & Bostock
Phlegmariurus schlimii (Herter) B.Øllg.
Phlegmariurus schmidtchenii (Hieron.) B.Øllg.
Phlegmariurus sellifolius (B.Øllg.) B.Øllg.
Phlegmariurus sellowianus (Herter) B.Øllg.
Phlegmariurus serpentiformis (Herter) B.Øllg.
Phlegmariurus setifolius (Alderw.) A.R.Field & Bostock
上思马尾杉 Phlegmariurus shangsiensis C.Y.Yang
Phlegmariurus sieberianus (Spring) B.Øllg.
鳞叶马尾杉 Phlegmariurus sieboldii (Miq.) Ching
Phlegmariurus silveirae (Herter ex Nessel & Hoehne) B.Øllg.
Phlegmariurus silverstonei B.Øllg.
Phlegmariurus sintenisii (Herter) B.Øllg.
Phlegmariurus societensis (J.W.Moore) A.R.Field
Phlegmariurus sooanus Lawalrée
Phlegmariurus sooianus Lawalrée
Phlegmariurus sotae (Rolleri) B.Øllg.
Phlegmariurus sphagnicola B.Øllg.
粗糙马尾杉 Phlegmariurus squarrosus (G.Forst.) Á.Löve & D.Löve
Phlegmariurus staudtii (Nessel) A.R.Field & Bostock
Phlegmariurus stemmermanniae A.C.Medeiros & Wagner
Phlegmariurus stephani B.Øllg.
Phlegmariurus strictus (Baker) A.R.Field & Bostock
Phlegmariurus subfalciformis (Alderw.) A.R.Field & Bostock
Phlegmariurus subintegrus (Hillebr.)
Phlegmariurus subtubulosus (Herter)
Phlegmariurus subulatus (Desv. ex Poir.) B.Øllg.
细叶马尾杉 Phlegmariurus subulifolius (Wall. ex Hook. & Grev.) S. R. Ghosh
Phlegmariurus tafaensis (Nessel) A.R.Field & C.W.Chen
Phlegmariurus tafanensis (Nessel)
Phlegmariurus talamancanus (B.Øllg.) B.Øllg.
Phlegmariurus talamauanus (Alderw.) A.R.Field & Bostock
Phlegmariurus talpiphilus (B.Øllg.) B.Øllg.
Phlegmariurus tardieuae (Herter) A.R.Field & Testo
Phlegmariurus tauri (Herter) A.R.Field & Testo
Phlegmariurus taxifolius (Sw.) Á.Löve & D.Löve
Phlegmariurus tenuicaulis (Underw. & F.E.Lloyd) B.Øllg.
Phlegmariurus tenuis (Humb. & Bonpl. ex Willd.) B.Øllg.
Phlegmariurus terrae-guilelmii (Herter) A.R.Field & Bostock
Phlegmariurus tetragonus (Hook. & Grev.) B.Øllg.
Phlegmariurus tetrastichoides (A.R.Field & Bostock) A.R.Field & Bostock
Phlegmariurus tetrastichus (Kunze) A.R.Field & Bostock
Phlegmariurus tico A.Rojas
Phlegmariurus toppingii (Herter) A.R.Field & Bostock
Phlegmariurus tournayanus Lawalrée
Phlegmariurus transilla (Sodiro ex Baker) B.Øllg.
Phlegmariurus treitubensis (Silveira) B.Øllg.
Phlegmariurus trifoliatus (Copel.) A.R.Field & Bostock
Phlegmariurus trigonus (C.Chr.) A.R.Field & Bostock
Phlegmariurus tryonorum B.Øllg.
Phlegmariurus tubulosus (Maxon) B.Øllg.
Phlegmariurus ulixis (Herter) B.Øllg.
Phlegmariurus unguiculatus (B.Øllg.) B.Øllg.
Phlegmariurus urbanii (Herter) B.Øllg.
Phlegmariurus vanuatuensis A.R.Field^[6]
Phlegmariurus varius (R.Br.) A.R.Field & Bostock
Phlegmariurus venezuelanicus (Herter) B.Øllg.
Phlegmariurus vernicosus (Hook. & Grev.) Á.Löve & D.Löve
Phlegmariurus verticillatus (L.fil.) A.R.Field & Testo
Phlegmariurus villonacensis (B.Øllg.) B.Øllg.
Phlegmariurus warneckei (Herter ex Nessel) A.R.Field & Testo
Phlegmariurus watsonianus (Maxon) B.Øllg.
Phlegmariurus weberbaueri (Nessel) B.Øllg.
Phlegmariurus weddellii (Herter) B.Øllg.
Phlegmariurus wilsonii (Underw. & F.E.Lloyd) B.Øllg.
Phlegmariurus xiphophyllus (Baker) A.R.Field & Bostock
云南马尾杉 Phlegmariurus yunnanensis Ching

杂交种：

Phlegmariurus ×koolauensis W.H.Wagner

圖示

展葉馬尾杉1
展葉馬尾杉2

引文出處

^ ^1.0 ^1.1 葉背隆起呈龍骨狀的覆葉馬尾杉.
^ ^2.0 ^2.1 垂枝石松. （原始内容存档于2019-03-16）. （页面存档备份，存于互联网档案馆）
^ Phlegmariurus (Herter) Holub. GBIF. [2023-02-15]. （原始内容存档于2023-02-15）. （页面存档备份，存于互联网档案馆）
^ ^4.0 ^4.1 ^4.2 ^4.3 ^4.4 Benjamin Øllgaard [B.Øllg.]. 2019. Synopsis of the Genus Phlegmariurus (Lycopodiaceae) in Colombia. Phytotaxa. 426(1); 1-105. DOI: 10.11646/phytotaxa.426.1.1
^ 江西武夷山新发现11种植物. NSII. [2023-02-15]. （原始内容存档于2023-02-15）. （页面存档备份，存于互联网档案馆）
^ Ashley Raymond Field. 2018. Phlegmariurus vanuatuensis (Huperzioideae, Lycopodiaceae) A New Species from Vanuatu, Re-circumscription of P. nummulariifolius and New Combinations in Phlegmariurus. PhytoKeys. 109: 53-66. DOI: 10.3897/phytokeys.109.29359

参考资料

台灣植物諮詢整合查詢系統-石松科（中文）.
特有生物研究保育中心台灣野生植物資料庫-垂直石松（中文）. ^{[失效連結]}
台中國立自然科學博物館-葉背隆起呈龍骨狀的覆葉馬尾杉（中文）.
台中國立自然科學博物館-姿態美麗的展葉馬尾杉（中文）.
台中國立自然科學博物館-垂枝石松. [2019-09-07]. （原始内容存档于2019-03-16）（中文）. （页面存档备份，存于互联网档案馆）
認識植物-垂枝石松（中文）.

[:0-1] 1.0 ^1.1 葉背隆起呈龍骨狀的覆葉馬尾杉.

[:1-2] 2.0 ^2.1 垂枝石松. （原始内容存档于2019-03-16）. （页面存档备份，存于互联网档案馆）

[3] Phlegmariurus (Herter) Holub. GBIF. [2023-02-15]. （原始内容存档于2023-02-15）. （页面存档备份，存于互联网档案馆）

[Ollgaard-4] 4.0 ^4.1 ^4.2 ^4.3 ^4.4 Benjamin Øllgaard [B.Øllg.]. 2019. Synopsis of the Genus Phlegmariurus (Lycopodiaceae) in Colombia. Phytotaxa. 426(1); 1-105. DOI: 10.11646/phytotaxa.426.1.1

[5] 江西武夷山新发现11种植物. NSII. [2023-02-15]. （原始内容存档于2023-02-15）. （页面存档备份，存于互联网档案馆）

[6] Ashley Raymond Field. 2018. Phlegmariurus vanuatuensis (Huperzioideae, Lycopodiaceae) A New Species from Vanuatu, Re-circumscription of P. nummulariifolius and New Combinations in Phlegmariurus. PhytoKeys. 109: 53-66. DOI: 10.3897/phytokeys.109.29359

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